Mineral Nutrition

All the living organisms have basic needs that are more or less similar. The macromolecules like fats, proteins and carbohydrates, and minerals and water are essential for their growth and development

First time in 1860, a German botanist, Julius von Sachs, showed that plants could be grown without soil in a nutrient solution. This technique of developing a plant in the nutrient solution is called hydroponics.

There are many nutrients that are essential for plants. Therefore, let us see what is the criteria that make them essential

Eligibility for essentiality

  • The element must be necessary to support the growth and reproduction of the plant.
  • In the absence of this element, the plants cannot complete their life cycle, and survival becomes very difficult.
  • The element must be irreplaceable and specific in function which means no other element can be used instead.
  • Its presence or absence affects the metabolism of the plant.

Macronutrients

These are generally present in tissues of plants in large amounts (more than 10mmole/kg of dry matter). Some of the macronutrients are –

  • Carbon
  • Hydrogen
  • Oxygen
  • Nitrogen
  • Phosphorus
  • Sulphur
  • Potassium
  • Calcium
  • Magnesium

While carbon, oxygen and hydrogen are obtained from water and carbon dioxide, other elements are obtained from the soil as mineral nutrition.

Micronutrients

These are also known as trace elements and are required in very small amounts (less than 10mmole/kg of dry matter). These include –

  • Iron
  • Magnesium
  • Manganese
  • Copper
  • Sodium
  • Silicon
  • Boron
  • Nickel

Essential elements can also be divided into four categories based on their various functions. These categories are:

  • Essential elements as an ingredient of biomolecules and hence structural elements of cells (e.g., carbon, oxygen, hydrogen and nitrogen).
  • Essential elements that are part of energy-related chemical compounds. (e.g., magnesium in chlorophyll and phosphorus in ATP).
  • Essential elements that stimulate or inhibit enzymes, for example, Mg2+ is an activator for both ribulose bisphosphate carboxylase oxygenase and phosphoenolpyruvate carboxylase that are notably significant enzymes in the process of photosynthetic carbon fixation. The zinc ion is an activator of alcohol dehydrogenase and Mo of nitrogenase.

Various forms and functions of essential nutrient elements are given below

Nitrogen

  • It is an essential nutrient required by plants in a large amount.
  • It is absorbed mostly as NO3, however, it can also be taken as NO2 or even NH4.
  • It is required by all parts of the plant, specifically for the growth of meristematic tissues and metabolically active cells.
  • It is one of the major ingredients of proteins, vitamins, nucleic acids and hormones

Phosphorus

  • Roots of the plant absorb phosphorus in the form of phosphate ions (either as H2PO4 or HPO4)
  • It is the main ingredient of cells membranes, some proteins, nucleotides and nucleic acids.
  • It is essential for all the phosphorylation reactions.

Potassium

  • It is absorbed by the plant as potassium ions.
  • It is required in large amounts in the buds, meristematic tissues, root tips and leaves.
  • It helps to regulate an anion-cation balance in cells and is involved in the synthesis of protein, opening and closing of stomata, maintenance of the turgidity of the cells and activation of enzymes.

Deficiency Symptoms of Essential Elements

The limitation in the supply of some essential element causes the retardation of growth. The concentration of these elements below a certain point, below which the plant starts showing retardation, is called critical concentration. The element is known to be deficient when it is below this critical concentration. Since every element has one or more specific function and role in the growth of the plant, its absence shows in the form of morphological changes. These changes signify the deficiency of a certain element and are called deficiency symptoms.

The deficiency symptoms are different for all elements. These symptoms disappear when the required element is provided to plant in the form of nutrition. However, if the element is not supplied to plant or is not available even after symptoms start showing, the plant may die. The symptoms are shown on the part of plant only where these elements can reach. Some elements are actively mobile in young developing tissues, and these plants show deficiency first in older tissues. For example – the deficiency of magnesium, potassium, and nitrogen can be first observed in senescent leaves. In the older leaves, biomolecules comprising of these elements are broken down that makes these elements available for mobilizing to younger leaves.

On the other hand, the relatively immobile elements tend to show the deficiency symptoms in the young tissues because these are not transferred out of the mature organs. For example, elements like calcium and sulphur form the structural part of the cell and hence are not easily released.

Some of the deficiency symptoms are chlorosis, stunted plant growth, necrosis, premature fall of leaves and buds, and restriction of cell division.

Chlorosis

  •  In chlorosis, chlorophyll is lost which results in yellowing of the leaves.
  • It is caused by the deficiency of elements like Fe, Zn, Mo, N, K, Mn, Mg and S.

Necrosis (death of tissue)

  •  It is mainly seen in leaf tissues.
  • It occurs due to deficiency of Mg, K, Cu, Ca.
  • It also occurs due to lack or low level of S, N, K and Mo which causes inhibition of cell division.

Delayed flowering is also seen in plants due to deficiency of N, S and Mo.

Absorption of minerals

Plants are capable of only absorbing the soluble minerals that can easily dissolve in water. The take minerals in the form of solution through the soil. From soil, these mineral are transported up through root hair cells. However, the soil is not a reliable source of mineral because the concentration of these minerals is very low.

Active transport

 

Water is transported in plants by the process of osmosis. However, mineral particles are more complicated than that and cannot be transported through osmosis. This is why active transport is used by plants for their movement.

The cells of the root hair have cell membranes which consist of carrier proteins. These proteins take the mineral ions and transport them across the membrane against the concentration gradient which means minerals do not move on the concept of higher to lower gradients. For this kind of transport, energy is required which is used in the form of ATP (adenosine triphosphate).